Shell injuries in Densepustula Lazarev, 1982, Pennsylvanian productidine brachiopod from the Donets Basin, Ukraine

Nine injured ventral valves of productidine brachiopods belonging to the genus Densepustula Lazarev, 1982, from the Mospyne, Smolyanynivka, Belaya Kalitva, and Kamensk formations (Upper Bashkirian-lower Moscovian, Lower and Middle Pennsylvanian) of the Donets Basin (eastern Ukraine) were studied. Three morphological types of damage traces have been recognized: Type A) longitudinal, sublongitudinal, and transversal thin straight or sinuous furrows, about 5-7 mm long and 0.5-1.5 mm thick; Type B) rounded and ellipsoidal pits, about 3-4 mm in diameter, located on the umbo, the anterior margin of the ventral valve, and the lateral slopes of the ventral valve; and Type C) irregularly-shaped dimples on the anterior margin and in the sulcus, 2-3 mm in size. These injuries are present on 9 out of 61 (15%) of ventral valves studied, but entirely absent from dorsal valves (0 out of 25). The most likely producers of these damage traces are cartilaginous fishes and parasites of unclear affinity.