BackgroundMost animal-pollinated flowers offer nectar or pollen as a reward to their pollinators.A small fraction, however, has switched to other rewards, such as fatty oils offered in glandular hairs or just below the epidermis.This type of reward has evolved in some 150 genera in at least 11 families, including South American and African Orchidaceae (Renner and Schaefer, 2010;Carneiro & Machado 2023).Oil-offering flowers are exploited by a few hundred species of oil-collecting bees in temperate and tropical
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