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This response addresses challenges and suggestions regarding the two-tiered life history model and its assumptions.

The two-tiered life history model: from interrogating assumptions to refining concepts and hypotheses

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This response addresses challenges and suggestions regarding the two-tiered life history model and its assumptions.
Reviewed commentator feedback on the two-tiered model.
Discussed proposed extensions and modifications of the model.
Explored the implications of varying levels of extrinsic mortality.
Showed broad support for the two-tiered model among commentators.
Identified key assumptions that require further clarification.
Highlighted the dualistic effects of extrinsic mortality on life history traits.

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Reply discusses challenges to the two-tiered life history model and its implications for child mortality and extrinsic mortality.

Ellis et al. (2025) studied this question.

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