Sea turtles from the Upper Cretaceous of the Penza Province (Russia)

The article describes remains of sea turtles (clade Pan-Chelonioidea) from three Upper Cretaceous localities of the Penza Province of Russia: Malaya Serdoba 1 and 2 (Campanian) and Penza (Maastrichtian). The distal part of the humerus of Pan-Chelonioidea indet. from the Malaya Serdoba 1 locality is similar to the humerus of Terlinguachelys fischbecki (Protostegidae) from the Campanian of USA. The turtle assemblage from the Malaya Serdoba 2 locality includes Protostega gigas (Protostegidae) and Pan-Chelonioidea indet. similar to T. fischbecki and representatives of the clade Ctenochelyidae, and resembles in systematic composition the sea turtle assemblage from the Campanian Beloe Ozero locality, Saratov Province of Russia. The outer surface of the costal plates of turtle from the Penza locality has a sculpture similar to Peritresius ornatus (Ctenochelyidae) and Glyptochelone suyckerbuycki.