R. C. Wilkinson, W. T. McClelland, Ruth M. Murillo, E. O. Ostmark, Stridulation and Behavior in Two Southeastern Ips Bark Beetles (Coleoptera: Scolytidae), The Florida Entomologist, Vol. 50, No. 3, 50th Anniversary (Sep., 1967), pp. 185-195
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Wilkinson et al. (1967) studied this question.