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James H. Peck, Carol J. Peck, Donald R. Farrar, Influences of Life History Attributes on Formation of Local and Distant Fern Populations, American Fern Journal, Vol. 80, No. 4, Symposium on Population Biology of Ferns (Oct. - Dec., 1990), pp. 126-142
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Peck et al. (1990) studied this question.
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