M. P. Harris, S. N. Freeman, S. Wanless, B. J. T. Morgan, C. V. Wernham, Factors Influencing the Survival of Puffins Fratercula arctica at a North Sea Colony over a 20-Year Period, Journal of Avian Biology, Vol. 28, No. 4 (Dec., 1997), pp. 287-295
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Harris et al. (1997) studied this question.
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