This paper lists over 100 species, mostly new records for Britain, found in the interstices of sand and shell gravel from North Wales beaches. The distribution of these species, other than the Kalyptorhynchia (Turbellaria) and the Gastrotricha, is given. The distribution of this fauna in relation to the substrate is discussed and species typical of fine-, medium- and coarse-particled substrates are listed.
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P. J. S. Boaden (1963) studied this question.
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