Peter G. L. Klinkhamer, Tom J. de Jong, Effects of Plant Size, Plant Density and Sex Differential Nectar Reward on Pollinator Visitation in the Protandrous Echium vulgare (Boraginaceae), Oikos, Vol. 57, No. 3 (Apr., 1990), pp. 399-405
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Klinkhamer et al. (1990) studied this question.
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