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Journal Article The Subspecies Concept and Its Taxonomic Application Get access E. O. Wilson, E. O. Wilson Search for other works by this author on: Oxford Academic PubMed Google Scholar W. L. Brown, Jr. W. L. Brown, Jr. Search for other works by this author on: Oxford Academic PubMed Google Scholar Systematic Biology, Volume 2, Issue 3, September 1953, Pages 97–111, https://doi.org/10.2307/2411818 Published: 01 September 1953
Wilson et al. (Tue,) studied this question.