Morris Goodman, John Czelusniak, G. William Moore, A. E. Romero-Herrera, Genji Matsuda, Fitting the Gene Lineage into its Species Lineage, a Parsimony Strategy Illustrated by Cladograms Constructed from Globin Sequences, Systematic Zoology, Vol. 28, No. 2 (Jun., 1979), pp. 132-163
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Goodman et al. (1979) studied this question.