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J. P. Grime, K. Thompson, R. Hunt, J. G. Hodgson, J. H. C. Cornelissen, I. H. Rorison, G. A. F. Hendry, T. W. Ashenden, A. P. Askew, S. R. Band, R. E. Booth, C. C. Bossard, B. D. Campbell, J. E. L. Cooper, A. W. Davison, P. L. Gupta, W. Hall, D. W. Hand, M. A. Hannah, S. H. Hillier, D. J. Hodkinson, A. Jalili, Z. Liu, J. M. L. Mackey, N. Matthews, M. A. Mowforth, A. M. Neal, R. J. Reader, K. Reiling, W. Ross-Fraser, R. E. Spencer, F. Sutton, D. E. Tasker, P. C. Thorpe, J. Whitehouse, Integrated Screening Validates Primary Axes of Specialisation in Plants, Oikos, Vol. 79, No. 2 (Jun., 1997), pp. 259-281
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Grime et al. (1997) studied this question.
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