John G. Lundberg, Maurice Kottelat, Gerald R. Smith, Melanie L. J. Stiassny, Anthony C. Gill, So Many Fishes, So Little Time: An Overview of Recent Ichthyological Discovery in Continental Waters, Annals of the Missouri Botanical Garden, Vol. 87, No. 1 (Winter, 2000), pp. 26-62
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Lundberg et al. (2000) studied this question.