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Masafumi Matsui, Takeshi Seto, Taeko Utsunomiya, Acoustic and Karyotypic Evidence for Specific Separation of Polypedates megacephalus from P. leucomystax, Journal of Herpetology, Vol. 20, No. 4 (Dec., 1986), pp. 483-489
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Matsui et al. (1986) studied this question.
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