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John P. Bryant, F. Stuart Chapin, III, David R. Klein, Carbon/Nutrient Balance of Boreal Plants in Relation to Vertebrate Herbivory, Oikos, Vol. 40, No. 3, Herbivore-Plant Interactions at Northern Latitudes. Proceedings of a Symposium Held 14-18 September, 1981, at Kevo, Finland (May, 1983), pp. 357-368
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Bryant et al. (1983) studied this question.
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