Key points are not available for this paper at this time.
Risto Kalliola, Jukka Salo, Maarit Puhakka, Marjut Rajasilta, New Site Formation and Colonizing Vegetation in Primary Succession on the Western Amazon Floodplains, Journal of Ecology, Vol. 79, No. 4 (Dec., 1991), pp. 877-901
Kalliola et al. (Sun,) studied this question.