A comprehensive analysis of threats to UNESCO WHSs in danger

• Threats to 54 WHSs in danger are examined. • WHSs in danger were distributed across 33 countries representing every region. • WHSs in danger share common threats. • The multidimensionality of the threats is assessed by clustering the threats into 13 primary factors. • Collaboration is identified as a key factor to promote the successful conservation and management of WHSs.