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David R. Anderson, Kenneth P. Burnham, Bruce C. Lubow, Len Thomas, Paul Stephen Corn, Philip A. Medica, Ronald W. Marlow, Field Trials of Line Transect Methods Applied to Estimation of Desert Tortoise Abundance, The Journal of Wildlife Management, Vol. 65, No. 3 (Jul., 2001), pp. 583-597
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Anderson et al. (2001) studied this question.
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