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Neil P. Schultes, Thomas P. Brutnell, Ashley Allen, Steven L. Dellaporta, Timothy Nelson, Jychian Chen, Leaf permease1 Gene of Maize Is Required for Chloroplast Development, The Plant Cell, Vol. 8, No. 3 (Mar., 1996), pp. 463-475
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Schultes et al. (1996) studied this question.
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