The accepted procedure for de termining production of multivoltine invertebrates by use of the Hynes method is to multiply the Hynes value by the number of generations per year. For aquatic insects, if pupal, adult, or egg stages comprise a significant portion of total generation time, this procedure will underestimate production. For crustaceans, if reproduction occurs before attaining the final size class, the procedure, using generation time, will overestimate production. It is necessary to multiply the Hynes value by 365/ CPI , where CPI is the cohort production interval (in days) from hatching to the attainment of the largest aquatic size class.
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Arthur C. Benke (1979) studied this question.
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