William B. Hopp, Studies on the Morphology and Life Cycle of Neoechinorhynchus emydis (Leidy), an Acanthocephalan Parasite of the Map Turtle, Graptemys geographica (Le Sueur), The Journal of Parasitology, Vol. 40, No. 3 (Jun., 1954), pp. 284-299
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W. Hopp (1954) studied this question.