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In a walking echidna the principal movement of the humerus is long-axis rotation. The humerus remains approximately perpendicular to the sagittal plane, but the femur is directed anterolaterally at angles from 35 degrees to 50 degrees . In addition to long-axis rotation, the femur elevates and depresses in an arc which usually varies between 40 degrees and 90 degrees . The femoral angle, the femoral elevation and depression, and the plantar contact of the manus beneath the glenoid are features found also in generalized therians.
Farish A. Jenkins (Fri,) studied this question.
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