J. W. Grula, T. J. Hall, J. A. Hunt, T. D. Giugni, G. J. Graham, E. H. Davidson, R. J. Britten, Sea Urchin DNA Sequence Variation and Reduced Interspecies Differences of the Less Variable DNA Sequences, Evolution, Vol. 36, No. 4 (Jul., 1982), pp. 665-676
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Grula et al. (1982) studied this question.