J. L. Harper, J. T. Williams, G. R. Sagar, The Behaviour of Seeds in Soil: I. The Heterogeneity of Soil surfaces and its Role in Determining the Establishment of Plants from Seed, Journal of Ecology, Vol. 53, No. 2 (Jul., 1965), pp. 273-286
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Harper et al. (1965) studied this question.
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