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The authors reconsider vesicle budding, this time including (besides bilayer bending elasticity) the energy of monolayer stretching, i.e. area dilation of one monolayer and compression of the other. Monolayer stretching during a shape change of the vesicle affects the spontaneous curvature of the membrane but, if measured values of material parameters are inserted in the equations, it turns out to be a minor perturbation. They also treat the budding of infinitely extended flat membranes where the spontaneous curvature is strictly constant during the deformation. The calculated shapes are compared to photographed buds in very large phosphatidylcholine membranes.
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Wiese et al. (1992) studied this question.
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