Journal Article On the Two Forms, or Dimorphic Condition, in the Species of Primula, and on their remarkable Sexual Relations Get access Charles Darwin, M.A., F.R.S., F.L.S., &c. Charles Darwin, M.A., F.R.S., F.L.S., &c. Search for other works by this author on: Oxford Academic Google Scholar Botanical Journal of the Linnean Society, Volume 6, Issue 22, March 1862, Pages 77–96, https://doi.org/10.1111/j.1095-8312.1862.tb01218.x Published: 06 January 2009
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Charles Darwin (1862) studied this question.