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The effect of wave exposure on the growth of low-shore populations of Perna perna (Linnaeus) was determined using 3 independent approaches at 6 sites on the south coast of South Africa: (1) Individual marking; notches were filed on the growing edges of mussels in situ. After 111 d, growth from the mark to the new growing edge was measured. (2) Internal shell-growth bands; mussels from the mark-recapture experiment were used to establish that growth bands are laid down tidally. Unmarked individuals were then used to measure growth over the last 60 tides.
McQuaid et al. (Sat,) studied this question.