Elizabeth L. Brainerd, Sheila N. Patek, Vertebral Column Morphology, C-Start Curvature, and the Evolution of Mechanical Defenses in Tetraodontiform Fishes, Copeia, Vol. 1998, No. 4 (Dec. 30, 1998), pp. 971-984
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Brainerd et al. (1998) studied this question.
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