Key points are not available for this paper at this time.
N. Jonsson, B. Jonsson, L. P. Hansen, Changes in Proximate Composition and Estimates of Energetic Costs During Upstream Migration and Spawning in Atlantic Salmon Salmo salar, Journal of Animal Ecology, Vol. 66, No. 3 (May, 1997), pp. 425-436
Jönsson et al. (Thu,) studied this question.