N. Jonsson, B. Jonsson, L. P. Hansen, Changes in Proximate Composition and Estimates of Energetic Costs During Upstream Migration and Spawning in Atlantic Salmon Salmo salar, Journal of Animal Ecology, Vol. 66, No. 3 (May, 1997), pp. 425-436
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Jönsson et al. (1997) studied this question.
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