Key points are not available for this paper at this time.
W. S. C. Gurney, R. M. Nisbet, J. H. Lawton, The Systematic Formulation of Tractable Single-Species Population Models Incorporating Age Structure, Journal of Animal Ecology, Vol. 52, No. 2 (Jun., 1983), pp. 479-495
Gurney et al. (Wed,) studied this question.