Anna M. Koltunow, Jessie Truettner, Kathleen H. Cox, Marco Wallroth, Robert B. Goldberg, Different Temporal and Spatial Gene Expression Patterns Occur during Anther Development, The Plant Cell, Vol. 2, No. 12 (Dec., 1990), pp. 1201-1224
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Koltunow et al. (1990) studied this question.