Steven F. Lehtinen, James B. Layzer, Reproductive Cycle of the Plains Minnow, Hybognathus placitus (Cyprinidae), in the Cimarron river, Oklahoma, The Southwestern Naturalist, Vol. 33, No. 1 (Mar. 30, 1988), pp. 27-33
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Lehtinen et al. (1988) studied this question.
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