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Sabo, J. 2003. Morris, W. F., and D. F. Doak. 2003. Quantitative Conservation Biology: Theory and Practice of Population Viability Analysis. Sinauer Associates, Sunderland, Massachusetts, USA. Conservation Ecology 7(2):2. https://doi.org/10.5751/ES-00512-070202
John L. Sabo (Wed,) studied this question.