Jerry R. Cox, M. H. Martin-R, F. A. Ibarra-F, J. H. Fourie, J. F. G. Rethman, D. G. Wilcox, The Influence of Climate and Soils on the Distribution of Four African Grasses, Journal of Range Management, Vol. 41, No. 2 (Mar., 1988), pp. 127-139
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Cox et al. (1988) studied this question.
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