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Florence A. Bernard, John M. Bernard, Patrick Denny, Flower Structure, Anatomy and Life History of Wolffia australiana (Benth.) den Hartog & van der Plas, Bulletin of the Torrey Botanical Club, Vol. 117, No. 1 (Jan. - Mar., 1990), pp. 18-26
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Bernard et al. (1990) studied this question.
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