The earthworm cuticle collagen was found to contain d-galactose, which constitutes about 12% of the total weight. By the alkaline borohydride treatment, followed by gel filtration, reduced di- and trisaccharides of d-galactose could be isolated. These oligosaccharides appeared to be linked to threonine and serine residues and account for about 70% of the total carbohydrate in the collagen.
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Lee et al. (1968) studied this question.
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