Richard D. Mace, John S. Waller, Timothy L. Manley, L. Jack Lyon, Hans Zuuring, Relationships Among Grizzly Bears, Roads and Habitat in the Swan Mountains Montana, Journal of Applied Ecology, Vol. 33, No. 6 (Dec., 1996), pp. 1395-1404
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Mace et al. (1996) studied this question.
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