David W. Coltman, Jill G. Pilkington, Judith A. Smith, Josephine M. Pemberton, Parasite-Mediated Selection against Inbred Soay Sheep in a Free-Living, Island Population, Evolution, Vol. 53, No. 4 (Aug., 1999), pp. 1259-1267
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Coltman et al. (1999) studied this question.