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The ichnospecies Trisulcus laqueatus is an epichnial trail that is constructed of three grooves separated by two ridges. It exhibits significant amounts of extramorphological variation because the maker was moving through wet sediment. Thus, most details of the maker are obscured. Nonetheless, there is evidence of it having been made by arthropods, including the sharply-angled turns in one example, the division of the lateral grooves into separate imprints in some locations along the length of the trail, and comparison to modern trails produced by Cydnidae (burrower bugs).
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Getty et al. (2022) studied this question.
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