B. D. Campbell, J. P. Grime, J. M. L. Mackey, A. Jalili, The Quest for a Mechanistic Understanding of Resource Competition in Plant Communities: The Role of Experiments, Functional Ecology, Vol. 5, No. 2, New Directions in Physiological Ecology (1991), pp. 241-253
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Campbell et al. (1991) studied this question.
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