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Lars O. Hedin, Joseph C. von Fischer, Nathaniel E. Ostrom, Brian P. Kennedy, Michael G. Brown, G. Philip Robertson, Thermodynamic Constraints on Nitrogen Transformations and Other Biogeochemical Processes at Soil-Stream Interfaces, Ecology, Vol. 79, No. 2 (Mar., 1998), pp. 684-703
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Hedin et al. (1998) studied this question.
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