D. S. Hik, R. L. Jefferies, A. R. E. Sinclair, Foraging by Geese, Isostatic Uplift and Asymmetry in the Development of Salt-Marsh Plant Communities, Journal of Ecology, Vol. 80, No. 3 (Sep., 1992), pp. 395-406
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Hik et al. (1992) studied this question.
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