E. van der Meijden, R. E. van der Waals-Kooi, The Population Ecology of Senecio Jacobaea in a Sand Dune System: I. Reproductive Strategy and the Biennial Habit, Journal of Ecology, Vol. 67, No. 1 (Mar., 1979), pp. 131-153
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Meijden et al. (1979) studied this question.
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