E. J. Chaneton, J. H. Lemcoff, R. S. Lavado, Nitrogen and Phosphorus Cycling in Grazed and Ungrazed Plots in a Temperate Subhumid Grassland in Argentina, Journal of Applied Ecology, Vol. 33, No. 2 (Apr., 1996), pp. 291-302
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Chaneton et al. (1996) studied this question.