J. Hugo Cota, Robert S. Wallace, Chloroplast DNA Evidence for Divergence in Ferocactus and its Relationships to North American Columnar Cacti (Cactaceae: Cactoideae), Systematic Botany, Vol. 22, No. 3 (Jul. - Sep., 1997), pp. 529-542
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Cota‐Sánchez et al. (1997) studied this question.
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