W. Armstrong, E. J. Wright, S. Lythe, T. J. Gaynard, Plant Zonation and the Effects of the Spring-Neap Tidal Cycle on Soil Aeration in a Humber Salt Marsh, Journal of Ecology, Vol. 73, No. 1 (Mar., 1985), pp. 323-339
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Armstrong et al. (1985) studied this question.
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