The hexaploid North American species Glandularia elegans has been crossed with the South American diploid species G. stellaroides, G. peruviana , and G. pulchella . Analysis of the behavior of chromosomes at meiosis in the hybrid indicates that the chromosomes of G. pulchella are similar to one of the genomes of G. elegans . Non‐homologous pairing and presence of multivalents also indicates that G. elegans is a segmental allohexaploid derived most likely from South American ancestors.
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Solbrig et al. (1968) studied this question.