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F. M. R. Hughes, The Ecology of African Floodplain Forests in Semi-Arid and Arid Zones: A Review, Journal of Biogeography, Vol. 15, No. 1, Biogeography and Development in the Humid Tropics (Jan., 1988), pp. 127-140
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Francine M. R. Hughes (1988) studied this question.
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