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John E. Fauth, Brian I. Crother, Joseph B. Slowinski, Elevational Patterns of Species Richness, Evenness, and Abundance of the Costa Rican Leaf-Litter Herpetofauna, Biotropica, Vol. 21, No. 2 (Jun., 1989), pp. 178-185
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Fauth et al. (1989) studied this question.
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