Bing-Quan Huang, William F. Sheridan, Embryo Sac Development in the Maize indeterminate gametophyte1 Mutant: Abnormal Nuclear Behavior and Defective Microtubule Organization, The Plant Cell, Vol. 8, No. 8 (Aug., 1996), pp. 1391-1407
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Huang et al. (1996) studied this question.